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Timothy
M. Evans
Associate Professor
Address:
Science Center 3017
Phone: (616) 395-7460
Email: evanst@hope.edu
Education
- 1989 - B.S. in Botany, University of Wyoming
- 1991 - M.S. in Botany, University of Wyoming
- 1995 - Ph.D. in Botany, University of
Wisconsin-Madison
Teaching Interests
- Cells and Genetics (BIO 240)
- Ecology and Evolution (BIO 280)
- Plant Systematics (BIO 343)
- Tropical/Island Botany and Geology (BIO 340; With
Dr. Bodenbender, Geological and Environmental Sciences): Includes Spring
Break trip to San
Salvador Island, Bahamas
- Senior Environmental Seminar (GES 401)
- Summer Flora (A five week field course in
beautiful northern Michigan at the Au Sable Institute)
Research/Scholarship
Interests

A maple tree
In front of me.
I wonder how
It came to be.
I'll make a tree
So I can see
The maple tree
Phylogeny.
(I don't work on
maples, but I couldn't get "Tradescantia" to fit my poem)
(Click here
for the rest of my poem)
My research involves
the use of both molecular (DNA sequence data and other molecular markers) and
morphological data to determine phylogenetic relationships among plants. Most
of my work involves members of the monocot families Commelinaceae (Wandering
Jew, Asiatic dayflower, etc.) and Bromeliaceae (pineapple family).
Within the
Commelinaceae, my students and I, in collaboration with Dr. Robert Faden
(Smithsonian Institution), are examining relationships within Aneilema,
Commelina, and the tribe Tradescantieae.
I am also
collaborating with Dr. Greg Brown (University of Wyoming) and Dr. Tania Wendt
(Universidade Federal do Rio de Janeiro, Brazil) in an NSF-funded
comprehensive study of phylogenetic relationships among members of subfamily
Bromelioideae (Bromeliaceae). As part of this project, I am exploring
the relationship between adaptive radiation and molecular evolution within
the subfamily. Additionally, this work is shedding new light on the
impacts that extinction rates may have on phylogenetic reconstruction.

Publications
de Sousa, L. O. F., T. Wendt, G. K. Brown, D. E. Tuthill, and T. M. Evans.
2007. Monophyly and phylogenetic relationships in Lymania (Bromeliaceae:
Bromelioideae) based on morphology and chloroplast DNA sequences. Systematic
Botany 32: 264-270
Givnish, T. J., J. C.
Pires, S. W. Graham, M. A. McPherson, L. M. Prince, T. B. Patterson, H. S.
Rai, E. H. Roalson, T. M. Evans, W. J. Hahn, K. C. Millam, A. W. Meerow, M.
Molvray, P. J. Kores, H. E. O'Brien, J. C. Hall, W. J. Kress, and K. J.
Sytsma. 2006. Phylogeny of the monocots based on the highly informative cpDNA
gene ndhF: evidence for widespread concerted convergence. Pp. 28-51, In: Proceedings
for the Third International Symposium on Monocots. Ontario, California.
Wade, D. J., T. M.
Evans, and R. B. Faden. 2006. Subtribal relationships in tribe Tradescantieae
(Commelinaceae) based on molecular and morphological data. Pp. 520-526,
In: Proceedings for the Third International Symposium on Monocots. Ontario,
California.
Givnish, T. J., J. C.
Pires, S. W. Graham, M. A. McPherson, L. M. Prince, T. B. Patterson, H. S.
Rai, E. H. Roalson, T. M. Evans, W. J. Hahn, K. C. Millam, A. W. Meerow, M.
Molvray, P. J. Kores, H. E. O'Brien, J. C. Hall, W. J. Kress, K. J. Sytsma.
2005. Repeated evolution of net venation and fleshy fruits among monocots in
shaded habitats confirms a priori predictions: evidence from an ndhF
phylogeny. Proceedings of the Royal Society London (Biology) 272: 1481-1490.
Givnish, T. J., K. C.
Millam, T. M. Evans, J. C. Hall, J. C. Pires, P. E. Berry, and K. J. Sytsma.
2004. Ancient vicariance or recent long-distance dispersal? Inferences
about phylogeny and South American-African disjunctions in Rapateaceae and
Bromeliaceae based on ndhF sequence data. Int. J. Plant Sci. (4 Suppl.):
S35-S54.
Evans, Timothy M.,
Robert B. Faden, Thomas J. Givnish, and Kenneth J. Sytsma. 2003.
Phylogenetic relationships in the plant family Commelinaceae. II:
A cladistic analysis of rbcL sequences and morphology. Systematic
Botany 28: 270-292.
Watson, L. E., P. L.
Bates, T. M. Evans, M. M Unwin and J. R. Estes. 2002. Molecular phylogeny of
Subtribe Artemisiinae (Asteraceae), including Artemisia and its allied and
segregate genera. BMC Evolutionary Biology 2:17
Evans, Timothy M.,
Robert B. Faden, Michael G. Simpson, and Kenneth J. Sytsma. 2000.
Phylogenetic relationships in the plant family Commelinaceae. I:
A cladistic analysis of morphological data. Systematic Botany 25:
668-691.
Evans, Timothy M., R.
B. Faden, and K. J. Sytsma. 2000. Homoplasy in the Commelinaceae: a
comparison of different classes of morphological characters. Pp. 547-556 in
Proceedings of the Second International Conference on the Comparative Biology
of the Monocots, Sydney, September 1998, eds. K. L. Wilson and D. A.
Morrison. Melbourne: CSIRO.
Givnish, T. J., T. M.
Evans, M. L. Zjhra, T. B. Patterson, P. E. Berry, K. J. Sytsma. 2000.
Molecular evolution, adaptive radiation, and geographic diversification in
the amphiatlantic family Rapateaceae: evidence from ndhF sequences and
morphology. Evolution 54(6): 1915-1937.
Watson, L. E., T. M.
Evans, and T. Boluarte. 2000. Molecular phylogeny of tribe
Anthemideae (Asteraceae), based on chloroplast gene ndhF. Molecular
Phylogenetics and Evolution 15: 59-69.
Faden, R. B. and T.
M. Evans. 1999. Reproductive characters, habitat and phylogeny in African
Commelinaceae. In Timberlake, J. & S. Kativu (eds.), African
Plants. Biodiversity, taxonomy and uses Proceedings of the 1997 AETFAT
Congress, Harare, Zimbabwe, pp. 23-38.
Givnish, T. J., T. M.
Evans, J. C. Pires, and K. J. Sytsma. 1999. Polyphyly and convergent
morphological evolution in Commelinales and Commelinidae: evidence from
rbcL sequence data. Molecular Phylogenetics and Evolution 12: 360-385.
Daugherty, John,
Timothy M. Evans, Tally Skillom, Linda E. Watson, and Nicholas P.
Money. 1998. Evolution of spore release mechanisms in the
Saprolegniaceae (Oomycetes): evidence from a phylogenetic analysis of
Internal Transcribed Spacer sequences. Fungal Genetics and Biology 24:
354-363.
Evans, Timothy
M. 1995. A phylogenetic analysis of the Commelinaceae based on
morphological and molecular data. Ph. D. Dissertation. University
of Wisconsin. Madison, Wisconsin.
Evans, Timothy
M. and Gregory K. Brown. 1990. Plicate staminal filaments
in Tillandsia subgenus Anoplophytum. J. Bromel. Soc. 40:
11-15.
Evans, Timothy
M. and Gregory K. Brown. 1989. Plicate staminal filaments
in Tillandsia subgenus Anoplophytum. Amer. J. Bot. 76: 1478-1485.
Evans, Timothy
M. and Gregory K. Brown. 1989. Stomata in Tillandsia
bryoides. J. Bromel. Soc. 39: 58-61.
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